1976 B.Sc.(Hons) Zoology and Applied Entomology, Imperial College, London 1977-78 Research Assistant, ROy Anderson and Phil Whitfield, Kings College, London 1978-82 D.Phil (Oxon) Zoology and Edward GRey Institute, Oxford University 1982-83 Post DOc with ROy Anderson Imperial College 1983-87 PostDoc with Bob May, Princeton, USA 1987-89 Assistant Professor, Biology, University of Rochester, New York 1990- Present ~ All forms of Professor, EEB, Princeton University 2011-Present External Faculty, Santa Fe Institute, New Mexico 2017-Present. A.D White Visting Chair, Cornell University, Ithaca, NY