My principal area of research is Evolutionary genetics and evolutionary ecology. I have a broad expertise in evolutionary biology, genetics and ecology. I have been working on adaptation and mutation, local adaptation, evolution of genetic systems, evolution of gene duplicates, speciation, genetic conflicts, dispersal, biotic interactions, statistics and fitness measures. I do theory but have been also working with many empirical systems (vertebrates, insects, crustaceans, fungi, plants, helminths, bacteria), in the lab and in the field.